By using DNA testing to match tusk pairs smuggled in separate consignments, Wasser and his team are able to link multiple ivory shipments to the same smugglers. Each map indicates separate shipments, with the location, date and weight of the seizure shown in the lower left. The blue circles show the geographic origins of the tusks based on genetic analysis; origins of a small number of poached elephant corpses matched to tusks are shown in open red circles. Linked pairs are shown by double-headed arrows, with thickness indicating the number of pairs. Shipments labeled "1" or "2" are linked by other lines of evidence.